Karolinska Institute (KI; Swedish: Karolinska Institutet)
Uppsala University (Swedish: Uppsala universitet)
Umeå University (Swedish: Umeå universitet; Ume Sami: Ubmeje Universitiähta)
Paris Sciences et Lettres University (PSL University or simply PSL)
Örebro University (Swedish: Örebro universitet)
Wageningen University & Research
